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Hymenoptera Holotypes of the Smithsonian Institution

Michael Gates Michael Gates

Matthew Buffington Matt Buffington


The arrangement of the specimen labels in the images on this site reflect their top to bottom positioning on the pin as viewed from upper left to lower right.

This website is a collaborative effort among several individuals. Michael Gates (USDA, Smithsonian Institution, Washington, DC) conceptualized the project. Robert Kula (USDA, Smithsonian Institution, Washington, DC) and Matthew Buffington (USDA, Smithsonian Institution, Washington, DC) contributed to the design of the website. Type photographs were taken by Mr. Jeffrey Chiu and Eugenio (Gino) Nearns (USDA APHIS PPQ NIS) developed the website and prepared the online searchable database.

Signiphoridae type specimens were photographed and processed by members of the Woolley Lab (Ana Dal Molin and Lydia Wessner) and support provided by National Science Foundation - PEET DEB-0730616 grant to Dr. Jim Woolley (Texas A&M University) and Dr. John Heraty (University of California, Riverside).

Additional Hymenoptera Types Databases
Smithsonian Institution Ant Type Specimen Image Database
Smithsonian Institution Bee Database


Robert Kula Robert Kula

Eugenio (Gino) Nearns Gino Nearns
Please email Michael Gates with error notification, suggestions, or comments.